We describe a striking new species of Grallaria antpitta from wet, upper su
btropical forest in the upper Rio Chinchipe drainage, provincia Zamora-Chin
chipe, Ecuador. Notes on its natural history and molecular systematics are
presented along with spectrograms of its voice. The species is readily diag
nosed by its large size, unique white facial markings, and voice. Initial r
esults from DNA sequence analyses place the new species in a well-supported
clade that includes Grallaria nuchalis (which is syntopic with the new spe
cies), G. hypoleuca, G. watkinsi, and G. ruficapilla.