The relief at the nest between the partners takes place twice a day: o
nce from female to male in the morning empty set = 4 h 11 +/-5 min (n
= 132) after sunrise and once from male to female in the late afternoo
n empty set = 2 h 06 +/-3 min (n = 160) before sunset. The first calls
to the relief are mostly given by males. During the relief ceremonies
males bow down their heads and front bodies more than females. By bow
ing they probably demonstrate their peaceful intention to the females.